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William Walker (guest star Tom Skerritt) dies unexpectedly at a family birthday celebration. Now his family will be forced to face some new life realities. Kitty's (Calista Flockhart) radio success has led to a TV job offer in Los Angeles, but her New York boyfriend has just proposed. Down to earth Thomas (aka Tommy) (Balthazar Getty) has joined forces with his sister, Sarah (Rachel Griffiths), in an effort to fix the family business, which is in deep financial trouble.

On the campus of Crandall College, three students live in the basement of Pi Nu fraternity, nicknamed "Le Dump". Rather than attend class, the three cellar-dwellers create havoc with the more straight-laced members of the fraternity, and interact with the Gamma Iota sorority sisters that live nearby.