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Koechner began studying improvisational comedy in Chicago at the ImprovOlympic, under the teachings of Del Close, before moving on to joining the Second City Northwest. After one-year stints on Saturday Night Live and Late Night with Conan O'Brien, he began taking small parts in films such as Man on the Moon and Wag the Dog. While filming the country mockumentary Dill Scallion, Koechner befriended Dave 'Gruber' Allen, forming The Naked Trucker & T-Bones Show. The act became a hit at Hollywood clubs such as Largo, and the duo were invited to open for Tenacious D.

His role as Champ Kind in Anchorman: The Legend of Ron Burgundy brought him many other films, including Talladega Nights, Waiting..., Tenacious D in the Pick of Destiny and Thank You for Smoking, in addition to recurring guest appearances on The 40 Year-Old Virgin co-star Steve Carell's The Office as Todd Packer. On January 17, 2007, Koechner's Naked Trucker & T-Bones Show premiered on Comedy Central, giving him a starring role and full creative control. David Koechner's first leading role as Coach Lambeau Fields in Fox Atomic's sports comedy, The Comebacks opened on October 19, 2007.

Koechner is married to actress Leigh Koechner and has four children, his 2 youngest are twins Sargent and Audrey, born June 2, 2006. He also performs regularly in Los Angeles at both the ImprovOlympic West theater (in a show entitled Beer Shark Mice), and at Largo, the nightclub where the Naked Trucker Show plays regularly. David Koechner supported the 2004 Presidential campaign of John Kerry, performing at a June 6, 2004 Hollywood fundraiser for the Senator, where The Naked Trucker & T-Bones opened up for Tenacious D. Koechner cites Lou Reed, Iggy Pop, Uncle Tupelo and its off-shoots (Wilco and Son Volt) as some of his musical favorites. He also strongly admires Creedence Clearwater Revival, whom he describes as, "the American Beatles."