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Stonestreet is perhaps most notable for his role as Ronnie Litre on CSI: Crime Scene Investigation (which he has portrayed 13 times as of February 10, 2005). He has also appeared on such television shows as Dharma & Greg, Malcolm in the Middle, Party of Five, Spin City, ER, The West Wing, Greg the Bunny, Providence, Close to Home, Crossing Jordan, and Bones. His first major film role came in 2000 when he played Sheldon the desk clerk in Almost Famous. He has also played Dr. Benson in Girls Will Be Girls, Ed the Trucker in The Island, and Courtney's Neighbor in Ninja Cheerleaders. In 2007, Stonestreet appeared in the short film Vinny's Vault which was produced during the reality show On the Lot.

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