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Weaving was born in Nigeria to English parents Anne, a tour guide, and Wallace Weaving, a seismologist. He spent his childhood in South Africa and then moved to the United Kingdom in his teens. While in England he attended the prestigious boarding school Queen Elizabeth's Hospital, commonly known in Bristol as QEH. He moved to Australia in 1976, where he attended Sydney's Knox Grammar School. Weaving later graduated from Australia's National Institute of Dramatic Art in 1981.

Weaving's first major role was in the 1984 Australian television series Bodyline, as the English cricket captain Douglas Jardine. Weaving appeared in the Australian miniseries The Dirtwater Dynasty in 1988 and starred opposite Nicole Kidman in the 1989 film Bangkok Hilton. In 1991, Weaving received the Australian Film Institute's award for "Best Actor" for his performance in the low-budget Proof. He also appeared as Sir John in the 1993 Yahoo Serious comedy, Reckless Kelly, a lampoon of the famous Australian outlaw Ned Kelly. However, Weaving first received attention overseas with the international hit Priscilla, Queen of the Desert in 1994. In 1998 Weaving received the award for "Best Actor" from the Montreal Film Festival for his performance in The Interview. Weaving was also a voice actor in the cartoon film The Magic Pudding.

He earned further international attention with his performance as the enigmatic Agent Smith in the 1999 blockbuster hit The Matrix. Weaving later reprised the role of Smith in that film's 2003 sequels: The Matrix Reloaded and The Matrix Revolutions.

He also garnered much popular attention in the role of the Elven Lord, Elrond in Peter Jackson's three-film adaptation of The Lord of the Rings, released between 2001 and 2003. Weaving was the main actor in Andrew Kotatko's award-winning film Everything Goes (2004). He also starred as a heroin addicted ex rugby league player in the 2005 Australian indie film Little Fish, opposite Cate Blanchett. He also played the title role as V in the 2006 film V for Vendetta, in which he was reunited with the Wachowski brothers, creators of The Matrix trilogy, who wrote the adapted screenplay. Actor James Purefoy was originally signed to play the role, but he pulled out six weeks into filming. Weaving appeared in the majority of V for Vendetta, and reshot all of James Purefoy's scenes as V (even though his face is never seen) apart from a couple of minor dialogue-free scenes early in the film. Stuntman David Leitch performed all of V's stunts.

Weaving also reprised his role as Elrond for the video game, The Lord of the Rings: The Battle for Middle-Earth II. He regularly appears in productions by the Sydney Theatre Company. In 2006, he worked with Cate Blanchett on a reprise of the STC production of Hedda Gabler in New York City. In a controversial move[citation needed] by director Michael Bay, Weaving was chosen as the voice of the Decepticon leader Megatron in the 2007 live-action film Transformers, rather than using the original voice created by the classic voice actor, Frank Welker. Bay stated on the DVD release of the film that he wanted Megatron to have a physicality similar to Weaving's, and that Welker's voice didn't fit the new interpretation of the character.

When he was 13 years old, Weaving was diagnosed with epilepsy. Due to the prospect of seizures, Weaving does not drive cars.[citation needed] He lives with his wife Katrina Greenwood and two children, Harry (b. 1989) and Holly (b. 1993). He has a brother, Simon Weaving who has two daughters, Samara Weaving and Morgan Weaving.

A notable pescatarian, Weaving is also the primary ambassador for Australian animal rights organization Voiceless. Hugo attends events and promotes Voiceless in interviews. He also assists Voiceless in their judging of annual grant's recipients.

• ...Maybe This Time 1980 Student 2
• The City's Edge 1983 Andy White
• Bodyline 1984 Douglas Jardine
• For Love Alone 1986 Johnathan Crow
• Melba 1987 Charles Armstrong
• The Right Hand Man 1987 Ned Devine
• Dadah Is Death 1988 Geoffrey Chambers
• The Dirtwater Dynasty 1988 Richard Eastwick
• Bangkok Hilton 1989 Richard Carlisle
• ...Almost 1990 Jake
• Proof 1991 Martin
• Road to Alice 1992 Morris
• Frauds 1993 Jonathan Wheats
• Reckless Kelly 1993 Sir John
• The Custodian 1993 Det. Church
• Seven Deadly Sins 1993 Lust
• Exile 1994 Innes
• The Adventures of Priscilla, Queen of the Desert 1994 Anthony 'Tick' Belrose/Mitzi Del Bra
• What's Going On, Frank? 1994 Strange Packer in Supermarket
• Bordertown 1995 Kenneth Pearson
• Babe 1995 Rex the Male Sheepdog
• Naked: Stories of Men 1996 Martin Furlong
• The Bite 1996 Jack Shannon
• True Love and Chaos 1997 Morris
• Halifax f.p: Isn't It Romantic 1997 Det. Sgt. Tom Hurkos
• Frontier 1997 Governor Arthur
• Babe: Pig in the City 1998 Rex the Male Sheepdog
• Bedrooms and Hallways 1998 Jeremy
• The Interview 1998 Eddie Rodney Fleming
• The Kiss 1998 Barry
• Strange Planet 1999 Steven
• Little Echo Lost 1999 Echo Man
• The Matrix 1999 Agent Smith
• The Lord of the Rings: The Fellowship of the Ring 2001 Elrond
• Russian Doll 2001 Harvey
• The Old Man Who Read Love Stories 2001 Rubicondo (Dentist)
• The Lord of the Rings: The Two Towers 2002 Elrond
• The Lord of the Rings: The Return of the King 2003 Elrond
• The Matrix Reloaded 2003 Agent Smith
• The Matrix Revolutions 2003 Agent Smith
• After the Deluge 2003 Martin Kirby
• Peaches 2004 Alan
• Everything Goes 2004 Ray
• Little Fish 2005 Lionel Dawson
• Happy Feet 2006 Noah
• V for Vendetta 2006 V
• The Key Man 2007 Vincent
• Transformers 2007 Megatron
• The Tender Hook 2008 McHeath
• The Wolf Man 2009 Detective Aberline

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