In the previous episode, Gul Dukat, disguised as a Bajoran named "Anjohl Tennan", got very close to Kai Winn, Benjamin Sisko married Kasidy Yates despite warnings from the Prophets, and Ezri Dax and Worf were beamed by the Breen aboard a Jem'Hadar ship.
Thot Gor, of the Breen, presents Dax and Worf to the changeling as a gift. The Breen are accepted into the Dominion regardless of Damar's disapproval of the treaty and its consequences for Cardassia. In addition, Septimus III is attacked by the Klingons and although Weyoun promised "deal with it", the entire Eleventh Order is wiped out. Damar's mood worsens as Thot Gor is allowed access to classified Cardassian information.
The mood in Quark's bar is sombre, as the crew misses Worf and Dax. Bashir's talk leads O'Brien to believe that he is in love with Ezri. Sisko isn't well either, as his wife begins to be uncomfortable with being married to an important figure in Bajoran religion.
During a confrontation with their captors, Worf slays Weyoun, though this is ultimately pointless as another clone would be sent in. As Worf and Ezri are taken to be executed, Damar realizes that the Dominion is not a good thing for Cardassia. He frees Worf and Dax and informs them he will help the Federation.
"Anjohl" convinces Winn to walk the path of the Pah-wraiths, by promising that Bajor will be "restored". However, Winn receives a vision from the Pah-wraiths that frightens her. She calls on Kira, who advises her to step down as Kai. Winn, disapproving of this advice, returns to Anjohl for comfort, and reluctantly accepts the Pah-wraiths.